Overview
- Distinctive dog skull morphology first appears around 11,000 years ago, aligning with genetic estimates for dog–wolf divergence.
- Early Holocene dogs already showed about half of today’s skull-shape variation, though extreme modern forms such as pugs and borzois were absent.
- The earliest clearly domestic specimen in the dataset comes from Veretye in northwest Russia (~10,800 years ago), with early dogs later identified in Illinois (~8,500 years ago) and Asia (~7,500 years ago).
- Late Pleistocene remains previously proposed as proto-dogs did not exhibit domesticated skull shapes, leaving the earliest domestication phase difficult to detect.
- A companion Science paper finds dogs traveled with migrating humans after the last glacial period and were sometimes traded between groups.